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Fish Reproduction

Animal Breeding

Pensacola Jr College

Female Fish Repro Tract

w    Ovaries compact and hollow

w    Eggs usually shed into ovarian cavity continuous with gonaduct (fish oviduct)

Ovary in Fish

Females - Specializations

w    Agnathans:

•    No oviduct

•    Eggs shed into coelom, then must travel to cloaca

w    Chondrichthyans:

•    Eggs fertilized and embryos develop in ovisac

Hagfish Ovary / tract

Shark Egg Case (mermaid’s purse)

Females - Specializations

w    Seahorses and pipefish:

•    Female has ovipositor (copulatory organ)

•    Inject eggs into male abdomen

w    Live bearers (eg. guppies):

•    Eggs fertilized and embryo develops in ovisac

Seahorses

Seahorse Eggs / Embryos

Pregnant Female Guppy w/ gravid patch

Male Fish Repro Tract

w    Sperm cells develop in association with Sertoli cells

w    Leydig cells produce androgenic steroid hormones

w    Sperm produced in:

•    Seminiferous tubules in bony fish

•    Spermatic ampullae (cavities) in sharks and agnathans

Testis types in Fish

Males - Specializations

w    Agnathans:

•    Mature sperm released directly into coelom

•    No sperm duct

•    Sperm exit through a pair of genital pores in abdominal wall

w    Gnathostomes:

•    Sperm ΰ epididymis ΰ sperm duct (carries urine too)

Male Hagfish – ducts drain kidneys only

Males - Specializations

w    Chondrichthyans:

•    Paired claspers for copulation

w    Live-bearing bony fish:

•    Gonopodium (modified anal fin) for copulation

w    Most bony fish:

•    External fertilization

•    Sperm expelled = “milt”

Shark Claspers

Gonopodium - guppy

Repro Tracts - Rays

Repro Tracts - Lungfish

Repro Tracts - Teleost Fishes

Fish Sex Determination

w    Genetics

w    Environment

Genetic Sex Determination

w    Heterogametic sex = produces 2 kinds of gametes and determines offspring sex

•    Usually male in fishes

w    Homogametic sex = produces one kind of gamete

•    Usually female in fishes

Environmental Sex Determination

w    Males and females may be distinct based on temperature etc.

w    Others often begin as females and become males later

w    Some hermaphroditic

Aquaculture and Fish Sex

w    Male fish usually more colorful, larger, etc.

w    Aquaculturists often attempt to produce all male populations

•    YY males are viable and produce all male offpring

w    Case study: Channel Catfish

Types of Egg-Layers

w    Egg scatterers

w    Egg-buryers

w    Egg-depositors

w    Mouth-brooders

w    Nest-builders

Egg scatterer: goldfish

Egg-buryer: killifish

Egg-depositor: cichlids

Mouth-brooder: betta

Nest-builder: gourami

Female Fish Reproductive Cycle

w    Oocyte development

•    Body size dependent

•    Not hormonal

w    Vitellogenesis

•    Longest period

•    Nutritionally dependent

Female Fish Cycle (cont’d)

w    Oocyte maturation

•    Water uptake

w    Spawning

•    Environment/courtship triggers

•    Can induce with carp pituitary extract or HCG

w    Recovery

•    Die or recover and return to vitellogenesis