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Parturition & Lactation

Animal Breeding

Pensacola Jr College

PARTURITION

Signs of Imminent Delivery

w    Tailhead prominence (due to pelvic ligaments relaxation)

w    Vulval swelling and mucus strings (by estrogen)

w    Behaviors: nesting, seclusion, etc (by prolactin)

w    Enlarged mammary glands

w    Calculate gestation period

Camel pelvic ligaments (72 hours to parturition)

Imminent parturition (ewe)

Mammary swelling (camel)

Mammary Swelling (mare)

Parturition

w    The birth process, cervical dialation through placenta delivery:

w    Step 1 = cervical dialation

w    Step 2 = uterine contractions / fetal expulsion

w    Step 3 = placenta expulsion

1. Cervical Dilation

w    Cervical canal relaxes and opens

w    Relaxin (a protein hormone):

•    Produced by ovary & placenta

•    Relaxes pelvic / cervical muscles

w    Fetus of most animals moves:

•    Forefeet pointed at cervix

•    Head between forelegs

 

Abnormal positioning

w    Breech = tail towards cervix

w    One or both feet turned back

w    Head turned back

Breech presentation

One leg back 

Hind legs only

Head back

Both forelegs back

Elbow lock

Twins - front and back

Four legs - one head

2. Fetal expulsion

w    Initial trigger = elevated fetal cortisol

w    Cortisol ΰ estrogens produced by placenta ΰ local transfer to uterus ΰ increased PGF2a ΰ local transfer to ovary ΰ CL regression ΰ decreased progesterone

2. Fetal expulsion (cont’d)

w    PGF2a ΰ small uterine contractions ΰ decreased blood flow to uterus ΰ anoxia ΰ increased fetal movement ΰ stimulates uterine nerves ΰ oxytocin release from post. Pituitary ΰ increased contractions (positive feedback loop)

Fetal expulsion (camel)

Fetal expulsion (camel)

Dystocia

w    Prolonged / difficult birth

w    Assistance required

w    Due to:

•    Breech or other problem in rotation

•    Twins

•    Large fetuses

•    Small pelvic areas

•    First pregnancy

3. Placental Expulsion

w    Stimulated by:

•    Secondary oxytocin surge

•    Low level uterine contractions

w    Retained placenta:

•    Due to twinning, high milk production, dystocia, or low vitamin A/E or selenium

•    Treatment is infusion of iodine and antibiotic solutions in uterus to prevent infections

Placental expulsion (camel)

Newborn after parturition

w    Changes in circulation

w    Oxygen enters through lungs, requires surfactant

w    Susceptible to temperature extremes

w    Slowly uses glycogen stores in liver

w    Needs colostrum (first milk)

 

Mother after parturition

w    Reproductive tract returns to normal:

•    Organs shrink to normal size

•    Recovery / repair of uterine endometrium

w    Postpartum anestrus = absence of cycling

w    Postpartum estrus = first estrus/ovulation after birthing

LACTATION

Mammary Gland Anatomy

w    Udder

w    Median and lateral suspensory ligaments

w    Teat:

•    Teat cistern

•    Gland cistern

•    Ducts

•    Lobes / lobules

•    Alveoli

Mammary Anatomy (cow)

Alveolus Anatomy

w    Lumen = milk storage area

w    Epithelial cells = produce milk

w    Capillaries = bring in vital nutrients for milk production

w    Myoepithelial cells = contract to cause milk to empty into ducts

Alveolus anatomy

Mammary Gland Development

w    Fetal mammary glands begin development due to:

•    Prolactin

•    Corticosteroids

•    Progesterone

w    Adult female in gestation:

•    Estrogen ΰ duct development

•    Progesterone ΰ gland dev

Lactation

w    Newborn suckling ΰ release of prolactin (or growth hormone)ΰ milk production

w    Young suckling ΰ sensory nerve stimulation ΰ oxytocin from posterior pituitary ΰ milk letdown (+ feedback)

Inhibition of Lactation

w    Stress ΰ cortisol release ΰ inhibits oxytocin

w    Excitement ΰ opiods in brain ΰ inhibits oxytocin

w    Either case results in less or no milk letdown

Milk Composition

w    Complete nutrition including:

•    Proteins

•    Carbohydrates

•    Fats

•    Water

•    Minerals

•    Vitamins

w    Composition varies greatly between species